Beerling, D. J., Osborne, C. P., & Chaloner, W. G. (2001). Evolution of leaf-form in land plants linked to atmospheric CO2 decline in the Late Palaeozoic era. Nature. 410: 352-354.
Dassler, C. L., & Farrar, D. R. (2001). Significance of gametophyte form in long-distance colonization by tropical, epiphytic ferns. Brittonia. 53: 352-369.
Kenrick, P., & Crane, P. R. (1997). The origin and early diversification of land plants a cladistic study. Washington, DC: Smithsonian Institution Press.
Knie, N., Fischer, S., Grewe, F., Polsakiewicz, M., & Knoop, V. (2015). Horsetails are the sister group to all other monilophytes and Marattiales are sister to leptosporangiate ferns. Molecular phylogenetics and evolution. 90: 140-149.
Mora, C. I., Driese, S. G., & Colarusso, L. A. (1996). Middle to late Paleozoic atmospheric CO2 levels from soil carbonate and organic matter. Science. 271: 1105.
Pinson, J. B., Chambers, S. M., Nitta, J. H., Kuo, L. Y., & Sessa, E. B. (2017). The Separation of Generations: Biology and Biogeography of Long-Lived Sporophyteless Fern Gametophytes. International Journal of Plant Sciences. 178: 1-18.
PPG 1: The Pteridophyte Phylogeny Group*. (2016) A community-derived classification for extant lycophytes and ferns. Journal of Systematics and Evolution. 54: 563-603. *This project was organized by E Schuettpelz, H Schneider, AR Smith, P Hovenkamp, J Prado, G Rouhan, A Salino, M Sundue, TE Almeida, B Parris, EB Sessa, AR Field, AL de Gasper, CJ Rothfels, MD Windham, M Lehnert, B Dauphin, A Ebihara, S Lehtonen, PB Schwartsburd, J Metzgar, L-B Zhang, L-Y Kuo, PJ Brownsey, M Kato, and MD Arana, with 68 additional contributors.
Rothfels, C. J., Li, F. W., Sigel, E. M., Huiet, L., Larsson, A., Burge, D. O., ... & Shaw, S. W. (2015). The evolutionary history of ferns inferred from 25 low-copy nuclear genes. American Journal of Botany. 102: 1089-1107.
Rothwell, G., & Stokey, R. (2008). Phylogeny and evolution of ferns: a paleontological perspective. In T. Ranker & C. Haufler (Eds.), Biology and Evolution of Ferns (pp. 332-366). Cambridge: Cambridge University Press.
Sato, T., & Sakai, A. (1981). Cold tolerance of gametophytes and sporophytes of some cool temperate ferns native to Hokkaido. Canadian Journal of Botany. 59: 604-608.
Schuettpelz, E., & Pryer, K. M. (2009). Evidence for a Cenozoic radiation of ferns in an angiosperm-dominated canopy. Proceedings of the National Academy of Sciences, 106: 11200-11205.
Sessa, E. B., Testo, W. L., & Watkins, J. E. (2016). On the widespread capacity for, and functional significance of, extreme inbreeding in ferns. New Phytologist. 211: 1108-1119.
Taylor, E. L., Taylor, T. N., & Krings, M. (2009). Paleobotany: the biology and evolution of fossil plants. London: Academic.
Testo, W., & Sundue, M. (2016). A 4000-species dataset provides new insight into the evolution of ferns. Molecular phylogenetics and evolution. 105: 200-211.
Watkins Jr, J. E., & Cardelús, C. (2009). Habitat differentiation of ferns in a lowland tropical rain forest. American Fern Journal. 99: 162-175.
Watkins, J. E., Mack, M. K., & Mulkey, S. S. (2007). Gametophyte ecology and demography of epiphytic and terrestrial tropical ferns. American Journal of Botany. 94:701-708.